Caulicolous or foliicolous. Conidiomata stromatic, variable from acervuloid and irregularly divided, to sessile or stalked shallow cups mimicking apothecia, scattered to gregarious, intraepidermal to intraperidermal in origin, erumpent, gelatinous; basal stroma and excipular tissue well developed. Conidiophores arising in the concavity of the conidioma, branched, septate, hyaline, smooth walled, branches terminating in conidiogenous cells and sterile hyphae, invested in mucus. Sterile hyphae simple or occasionally irregularly branched in the apical part, straight or irregularly coiled at the apex, septate, hyaline, smooth walled. Conidiogenous cells phialides, cylindrical or subcylindrical, hyaline, smooth walled, proliferating percurrently up to three times. Conidia blastic-phialidec, four or five radiate, hyaline, smooth walled; main axis short, clavate to narrow obconic or subcylindrical with a narrow truncate base bearing minute marginal frills, zero or one septate; arms three or four, divergent, attenuated, septate, guttulate, smooth walled, unequal; after secession of conidium, base of the main axis often drawn out into a narrow, protuberant appendage of varying length.
TYPE ANAMORPH SPECIES: Crucellisporiopsis gelatinosus Nag Raj

Caulicola vel foliicola. Conidiomata stromatica, acervuloidea et irregulatim divisa vel cupuaas sessiles ad pedicellatas apothecia similia, dissita vel gregaria, originem intra-epidermalia vel intra-epidermalia, erumpentia, gelatinosa; stroma basali et tela excipuli bene evoluta. Conidiophora cavitatem conidiomatis enascentia, ramosa, septata, hyalina, laevia, ramuli in cellulis conidiogenis et hyphis sterilibus terminantes, muco involuta. Hyphae steriles simplices vel partem apicalem irregulatim ramosae, apicem rectae vel irregulatim torsivae, septatae, hyalinae, laeviae. Cellulae conidiogenae phialides, cylindraceae vel subcylindraceae, hyalinae, laeviae, usque ad ter percurrently prolificantes. Conidia blastico-phialidica, 4-5 radiata, hyalina, laevia; axis principalis brevis, clavatus vel anguste obconicus vel subcylindraceus, basi anguste truncatis cum minutis fimbris marginalibus, 0-1 septatis; brachia 3-4, divergentia, attenuata, septata, guttulata, inaequalia; in conidia liberata, basis axis principalis saepe in unam, longitudinum differentem, protrudentem appendicem producta.

Two anamorphs, one on decaying leaves of Weinmannia racemosa from New Zealand and the other on unidentified twigs of a tree from Venezuela, appeared superficially similar to Eriosporella calami (Niessl) Höhn. Both collections possess tetraradiate conidia (Figs. 4, 13) and percurrently proliferating phialides (Figs. 8, 12) borne in stromatic conidiomata. The anamorph from New Zealand has acervuloid to shallow cupulate, but gelatinous, conidiomata (Fig. 9) appearing as yellowish to orange yellow erumpent crusts. The conidiomata of the anamorph from Venezuela, on the other hand, mimic apothecia (Figs. 1, 2), the true nature of the fungus becoming apparent only after close scrutiny (Figs. 3-8). Both anamorphs have septate sterile elements which arise as branches of the conidiophores. In the New Zealand collection, they are coiled at the apices, but remain straight in the Venezuelan specimen.
Of the coelomycetes with tetraradiate conidia, only two anamorph genera need to be considered for comparison with the unnamed collections mentioned above. Eriosporella was proposed by Höhne1 (1916) to accommodate Cryptosporium calami Niessl. According to Sutton (1980), E. calami has eustromatic condiomata consisting of a central locule, a thin basal wall, and a thicker lateral wall extending into the disorganized epidermal tissue around the conidioma; walls composed of hyaline, thin-walled textura angularis to textura intricata; hyaline, branched, septate conidiophores; and phialidic conidiogenous cells with minute apertures and apically thickened periclinal walls, but without collarettes (cf. Fig. 267B, Sutton 1980). Although Sutton makes no mention of sterile elements, his illustration of conidia inside the conidioma (Sutton 1980, Fig. 266B) could be interpreted as representing such elements. An illustration of E. calami prepared from an isotype specimen in DAOM by Nag Raj and DiCosmo (1980) does not agree with Sutton's account of the fungus. The conidiogenous cells are actually annellides and the illustration given by these authors confirms that the conidiomata of E. calami lack sterile elements. Crucellisporium Farr has conidiomata with sterile elements, and tetraradiate conidia, but can be ruled out as a possible taxonomic niche for the two unnamed congeneric collections by its holoblastic-sympodial conidium ontogeny (Nag Raj 1974; Nag Raj and Kendrick 1978). A new anamorph genus is, therefore, necessitated.